There are unexpected outcomes for both families when an infestation at the nursery burrow forces the Whiskers to make a risky move with their youngest pups. Meanwhile, heavily-pregnant Flint Lockwood leads Hakuna Matata in search of a new home.

All three families of the dynasty are suffering a hard winter. Swift's life is threatened once again only this time the enemy is within. Flint is forced to take risks to feed her family and Daisy takes on the Zulus.