Domenico takes information he's extracted from Satu to menacing vampire Gerbert, who gains the advantage he has been waiting for. After sending Juliette into the world to do his bidding, Gerbert consults with Meridiana, the witch whom he keeps in thrall, but what does the prophecy she delivers mean? In Oxford, Diana refuses to help Knox realise his supremacist longing to destroy the vampires and answers his threats with such a violent force of untamed elemental power, as to set in motion a wider chain of events that bring her closer to Matthew but place her in even greater jeopardy.

Gillian pays a heavy price for her disloyalty to Diana, but not before news that vampires are studying witch DNA reaches Knox. The fall-out threatens the fragile equilibrium between creatures and the focus is on ending Matthew and Diana's relationship whatever the cost. Agatha treads a careful line of democracy at the Congregation, but Nathaniel continues to instigate daemon direct-action. Ysabeau relies on shock tactics to expel Diana until Matthew returns with breathtaking news about Diana's DNA. Diana and Matthew are finally united in love, only for the unexpected to happen