Hugh Dennis and the team visit Studley Royal Park in North Yorkshire, on a quest to find the lost Georgian mansion that would have been at the heart of the world class Water Gardens, which still exist today. The house and gardens were created by John Aislabie, but his house has vanished from the landscape and the only clue the team has to go on is a fallen tree that's revealed some mysterious bricks. With the help of the local community, the team uncovers the foundations of this Georgian mansion and finds out about the grand lives of the family who once lived there.

Hugh Dennis and the team visit the village of Strelley in Nottinghamshire, on a quest to find the long-lost medieval castle of a knight from the 14th century. Sampson de Strelley came from a long line of knights and rose to become the Sheriff of Nottinghamshire, but the castle he built has vanished from the landscape.