Will forms the Acafellas, an all-male acapella vocal group, and spends more time building his own confidence than he does with the glee club. In Will's absence, the glee club decides to hire a well-known choreographer to help coach them to Nationals. Meanwhile, Mercedes is bitten by the love bug, but her feelings aren't reciprocated. When the Acafellas perform for the P.T.A., they get a surprise visit backstage from Josh Groban.

Will thinks the glee club desperately needs more of an edge, so he brings in his former classmate, April Rhodes, to spice things up. Will's plan seems to be going smoothly until he realizes he may be losing a key member of the club.