As Brianna grapples with the life-changing revelations of the past summer, Claire must help her come to terms with the fact that she is truly her father's daughter – her 18th century Highlander father. To complicate matters further, Roger brings news that forces Claire and Brianna to face an impossible choice.

In the aftermath of a violent confrontation, Claire follows her conscience as a surgeon, even though it could put her and Jamie's lives at risk. At the same time, Jamie attempts to evade the reach of the Crown as its representative closes in on his illegal dealings.