The guys are invited to a wedding of a mutual friend. Nick, fearful he will run into his ex-girlfriend Caroline, asks Jess to be his date. Meanwhile, Winston takes his role as an usher very seriously and gets competitive on the dance floor with a young guest. Schmidt not only catches the eye of a beautiful woman, but also his "frenemy" Gretchen whom he sleeps with at every wedding reception.

Cece convinces Jess that Nick likes her more than just as a roommate and Schmidt tries to seal the deal with Cece.