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Defiance is under siege. As Nolan, Amanda, Irisa and the militia defend the town against the Votanis Collective, Datak recalls his father's chilling lessons and makes a pivotal decision, while Stahma takes an unexpected trip.
A visiting arms dealer changes Defiance's fortunes, and divides Amanda, Nolan and Berlin; at the same time, Nolan encounters difficulty recruiting a militia to fight the oncoming Votanis Collective army. Elsewhere, Stahma's presence causes strife among the mysterious Omec; and Datak and Alak have a reckoning.
A gruesome murder is committed by an unusual beast that's on the loose and being hunted down by Nolan and Irisa, while Doc Yewll delves into the surprising victim. Elsewhere, Amanda resists welcoming back Stahma; and T'evgin learns an unsettling truth about his daughter.