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The 2nd Mass encounters new people as Tom happens upon his former mentor, yet trust issues abound. Meanwhile, Maggie suspects that Pope will soon exit from the group; and Tector embraces his military past.
The unit marches onward to Charleston, S.C., as some in the group continue to grieve recent losses. Meanwhile, information surfaces about Tector's past; and Matt connects with a recently discovered harnessed girl.
In the Season 2 finale, the skitters draw nearer to the newly consolidated resistance force. Elsewhere, the 2nd Mass encounter their most perilous threat as Tom discovers game-changing information.
The season's plot focuses on the 2nd Massachusetts' discovery that a large community of survivors has formed in Charleston, their journey there, and their reception once they arrive. It focuses on the discovery that the Skitters are themselves "harnessed" and mind-controlled by the invaders, but that some of them are resistant to the effects and are rebelling against their Overlords.