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Tom's life hangs in the balance as his beamer drifts into space; Tom encounters a surprising guest; Anne reluctantly steps into Tom's shoes to help lead the 2nd Mass.
A surprise skitter attack leaves the 2nd Mass without food. In desperation, Maggie, Ben, Pope, and Sara go on a dangerous mission hoping to find supplies. Along the way, they encounter a family deeply scarred by the Espheni plan. Maggie must make false promises to insure the 2nd Masses' survival. Meanwhile, Tom and Anne discover the Espheni have created something part Espheni, part Skitter, part Hornet.. and part human.
Tom and the 2nd Mass are filled with rage, ready to destroy the enemy in an all-out battle to determine the fate of Earth.