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M.E.C.H. capture Breakdown to take him apart and implement the gathered knowledge into their secret plans. The Autobots thus have to rescue their own enemy.
Jack, Raf and Miko get trapped in-between dimensions, along with the revived body of the fallen Decepticon Skyquake. The Autobots have to figure out a way to save them. The battered and humiliated Starscream meanwhile tries to follow up on his latest scheme to overthrow Megatron.
M.E.C.H. and Airachnid team up to kidnap Jack's mom, and Arcee has to offer herself as ransom. M.E.C.H. is after the Transformer technology, while the sadistic Airachnid simply wants to kill everyone involved.