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Following a group of Manda, a giant monster makes landfall in Tokyo. Red dust forces Mei and Professor Li's flight to London to divert to Milan.
Mei discovers a mysterious code in Professor Ashihara's research. The Otaki crew confront Anguirus as part of the city's extermination efforts.
Ground forces under the command of Matsubara respond to Godzilla's attack on Tokyo. Now in London, Mei attempts to decipher Ashihara's research.