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The evil genius Karl sets out to destroy King Julien. When both Clover and Maurice are captured, King Julien must save them from his new nemesis.
King Julien learns that 99% of the lemurs love him, so he makes it his mission to find and convert his one "hater."
Exhausted by settling his lemur subject's disagreements, King Julien replaces himself with a dummy king so he can play hooky, unwittingly sparking a rebellion.
Prince Julien becomes king of the lemurs, but he quickly discovers that he has to do more than throw parties if he's going to be a successful leader.