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When Maurice learns that he isn't a lemur like everyone else, he sets off on a journey to find his Aye-Aye parents and sister Brosalind.
Julien is lost in the jungle and accidentally saves a Mountain Lemur named Koto who turns out to be the biggest threat the kingdom has ever faced.
When the kingdom is overrun with refugees, Julien is shocked that an old friend Koto and his Mountain Lemur army are preparing to attack.
King Julien and his trusty advisers race to save the kingdom from secret agents, stampeding clones and more -- while keeping the party going strong.