Team Gurren's journey brings them to an underground village, where society mandates a humbling existence and the worship of Gunmen as divine beings. Team Gurren is joined by Rossiu and two children, Gimmy and Darry. 

One of the Four Divine Generals of the Spiral King arrives to support Viral's Enki. Seizing the opportunity to land a great victory, Kamina and Simon pilot the Gurren-Lagann against seemingly overwhelming odds.