Yoko had been working on an island as a teacher until she learned of the situation at Teppelin and returned to break Simon free. Reunited, Simon and the members of Team Dai-Gurren launch themselves into space to protect the Arc-Gurren from the enemy forces.

The members of Team Dai-Gurren make their preparations for the imminent war against the Anti-Spirals, but Rossiu is missing. However, when Kinon tells Simon about what he really intends to do, both begin desperately searching for him.