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接上集,博士身受重伤,被迫重生,但在重生时博士利用了圣诞节被砍下的手(详见2005年圣诞特别篇和S3E11)重生并保持了原样。
就在这时躲在塔迪斯中的多娜在博士之手的召唤下触碰它,砍下的手又重新长出一个博士,多娜也拥有了半个时间领主的力量...罗斯和她的母亲以及新生的人类博士被送到恶狼湾,博士称担心他的分身会被报复,希望罗斯能照顾他,并对她说他就是我。
多娜希望留在博士身边却因无法处理博士那庞大的数据和记忆,被博士清除记忆后送回家过正常人的生活...
博士匆忙回到地球,一切正常,然而当博士进一步侦查时,怪事出现了,塔迪斯留在原地,而地球却不见了...于此同时,地球上的人类发现天空飘满了星球和宇宙飞船。
博士来到影子联盟(银河系警察总局),查询到共有27颗星球丢失。由于地球被放在了美杜莎时空瀑布中,博士找不到地球。
戴立克轮番轰炸城市,控制地球政府、交通,带走人质进行试验。在危机关头火炬木、莎拉简等人的努力下将博士拉到了地球同步时空中。刚出塔迪斯的博士,看到了罗斯,但随后被戴立克击中,被迫重生。
【博士第11次重生】
The beginning of the episode depicts the Tenth Doctor composing Ode to the Universe: a symphony based on the "music of the spheres"—an aural representation of the Universe's gravity patterns. During the composition, a Graske teleports into the TARDIS to warn the Doctor about the imminent opening of a portal within the TARDIS. The Doctor approaches the portal, linking the TARDIS to the Doctor Who Prom at the Royal Albert Hall. He briefly converses with the audience and then conducts the orchestra in a performance before he realises the Graske has escaped into the Prom with his water pistol. He forces the Graske's return by "reversing the polarity of the neutron flow" and banishes him from the TARDIS, to the other side of the galaxy. At the end of the episode, he tells the viewer that the music of the spheres encompasses everyone, and that "Everyone's got a song inside them. Every single one of you."