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Amid rising Welsh nationalism in the run-up to his investiture as Prince of Wales, Charles is sent to spend a semester there and learn the language.
Pursuing improvements in her racehorse management, Elizabeth imagines the life she might have had. Meanwhile, a plot against her government is brewing.
As the first men land on the moon, middle-aged Philip feels dissatisfied by his lack of achievement and begins searching for inspiration.
As the 1960s and '70s unfold, the royal family contends with conflict and betrayal while striving to uphold tradition in the face of an evolving world.