The great migration leads to a series of confrontations for the families on the great savannah. Tragedy strikes for lioness Kali, forcing her to make a difficult decision. Bakari challenges the jealous leader of the baboon troop, leading to a violent battle, and elephant mother Nalla has to decide the future of her badly behaved teenager Tembo. Shani the zebra mother has no choice but to cross the river, fully aware a crocodile is waiting - and if he does not get her, the currents might well sweep her away.

The aftermath of the devastating fire has left a blackened smoking wasteland, but rain soon brings hope to the scorched earth. Kike the cheetah and wild dog leader Jasari are both teaching their cubs how to survive on their own, only to be interrupted by a grieving giraffe. A baby elephant struggles in the flooded river, while Bakari returns when his family are threatened by a rival baboon troop. Lioness Kali has surprises in store which bring hope for the future of her pride, and as things draw to a close, a death brings all the families together in a remarkable truce.