Police Captain William Boone is caught between two worlds when he is recruited by the leader of the Talons, who are also known as the 'Companions' after saving his life at a public event. Boone becomes the Commander of Security and Interspecies Relations and is then given a opportunity to join the human resistance movement.

A young woman becomes a pawn in a power play between the companions and a charismatic religious leader when the Talon's use a experimental process to regenerate her hands.