When a hostile alien ship fires first at the Moya, the crew, despite the fact that their energy shield is up, finds their minds switched into another person's body. As a result, Crichton is in the body of Aeryn, Aeryn is in the body of Rygel, and Rygel is in the body of Crichton. In the cockpit, Pilot is in the body of Chiana, Chiana is in the body of D'Argo, and D'Argo is in the body of Pilot. They're all forced to adapt to their new bodies as they attempt to figure out what caused the switch.

Moya arrives at a planet where the people test their genetic compatibility by kissing. Crichton kisses a princess, who is next in line for the throne, and discovers that his DNA is a perfect match for her. He must now marry her and they will rule together. This is against the wishes of her brother, who is second in line for the throne -- conspiring with a Scarran.