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To protect a volatile pop star from an obsessed fan, Echo goes undercover as her new backup singer. Meanwhile, Agent Ballard is led down a dangerously wrong path in his investigation, and Lubov harbors a secret.
Echo is imprinted to be the female counterpart to an avid outdoorsman and quickly learns how the hunter becomes the hunted during her wilderness adventure. Agent Ballard receives a clue to Echo's real identity, encouraging him to continue his investigation, while flashbacks reveal the tragic events at the Dollhouse that led to Boyd's arrival and Dr. Saunders' scars.
Echo's stint as a midwife doesn't prove as laborious as when she becomes an expert safe-cracker, and more than an invaluable piece of art is stolen from her on the engagement, forcing Sierra to take on a familiar persona. Agent Ballard gives Lubov an ultimatum, and Adelle confides shocking information to Topher.
The Dollhouse is a very secret, and very illegal, place where wishes come true. Clients with the right connections and enough money can hire "Actives", people who have been programmed to perfectly fulfill the needs, and desires of their clients. The Actives are people who have chosen, for their own reasons, to surrender their bodies and minds for a five-year stay in the Dollhouse. Now they can be imprinted with any personality, skill, or even muscle memory. They can be the perfect companion, lover, spy, assassin; and when the job is done they forget everything. But something is wrong with the Active called "Echo". No longer just a blank slate waiting for her next assignment, Echo is remembering flashes of the lives that she has lived and the games she has played, and she is starting to wonder just who she really is.