The Doctor teaches Seven Of Nine about human behavior and how humans interact with each other while dating. The Doctor learns a bit about his own possible humanity during the lessons. After her disastrous first date with a crew member, the Doctor personally guides her, but he soon realizes he is falling in love with her. Meanwhile, Neelix guides an ambassador of a monastic colony through the ship, unwittingly creating a potentially embarrassing situation for the ambassador when he requests forbidden food and drink.

When Voyager is destroyed, Captain Braxton of the 29th Century Timeship Relativity recruits Seven of Nine to travel back in time and find who planted the "temporal disruptor." However, to save Voyager, she must do this without being discovered by the past Janeway.