On DS9, Sisko agonizes over his vision from the Prophets, warning him not to marry Kasidy, and Kai Winn receives a vision foretelling the arrival of a guide who will help her restore Bajor. Meanwhile, Ezri and Worf are interrogated aboard a Breen ship and unexpectedly witness a disturbing turn of events, one which may change the course of the entire war.

The Breen-Dominion Alliance launches a devasting attack at the heart of the Federation, and the Federation Alliance struggles to maintain its foothold in Cardassian space. Kai Winn, under the guidance of Dukat, begins researching forbidden texts about the Pah-wraiths. On Cardassia, Damar begins a rebellion against the Dominion.