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Despite the growing tension between them, Cole and Cassie travel to 1940s New York during World War II to track down a dangerously seductive member of the Twelve who's plotting an assassination.
When an attempt to destroy the virus changes the future, Cole finds himself stranded in 2016 with an unstable Jennifer Goines, who may or may not be his key to getting home. Meanwhile, Jones and Railly must deal with both the consequences of the changed timeline and the temporal facility's newest prisoner -- Ramse.
Jones reluctantly places her trust in Ramse to bring Cole and Cassie safely home after they get stranded in the year 1944. As Ramse looks forward to this chance to atone for his sins, Jones explores the true nature of time with old Jennifer Goines.
Cole and Cassie must travel to different time periods to stop The Army of the 12 Monkeys, who seek to unravel time itself. Plus: The identity of the leader of the Army, the Witness, is revealed.