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While searching for Jennifer Goines in post-WWI France, Cole and Jones encounter a dangerous group of time-travelers‒the Guardians.
In the apocalypse of 2163, a desperate Cole searches for Titan. Meanwhile, a captive Railly is faced with an impossible choice as the 12 Monkeys await the arrival of The Witness.
Bonds of trust are tested when Ramse returns to the facility with an old enemy held captive. How far will Cole and Jones go for the answers they need?
With the identity of the Witness - the engineer of the reality ending Army of the 12 Monkeys - now known, Cole and his allies must travel through different eras in history to stop him.