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While Albizzi uses the plague to turn Florentines against Cosimo and his dome, two Medici family members are lost. Marco forces a shocking confession.
As the war with Milan takes a heavy toll on Florence, Cosimo boosts morale and the economy by undertaking the impossible: building the cathedral dome.
After Cosimo's arrest, his wife tries to buy a Signoria member's help. Piero seeks to refute the charges, whilst Lorenzo takes a more drastic approach.
After his father's murder, banking heir Cosimo Medici battles opponents of his artistic, economic and political visions for 15th-century Florence.