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After 20 years of grooming, Cosimo Medici replaces his late father in Florence's Signoria, where he must decide whether to support a call for war.
As the war with Milan takes a heavy toll on Florence, Cosimo boosts morale and the economy by undertaking the impossible: building the cathedral dome.
After his father's murder, banking heir Cosimo Medici battles opponents of his artistic, economic and political visions for 15th-century Florence.