To speed along the investigation into her house fire, Betty is pressured to date an annoying fireman. Wilhemina tries to prove to Don, an old flame from her past, that she's just "Wanda from the block" instead of an uber-bitch, and Amanda draws closer to Tyler, convincing Claire to hire him as an in-house model at Mode. Meanwhile, Hilda meets Bobby's parents for the first time.

Betty is thrilled to get her braces off, courtesy of her chatty new orthodontist, Dr. Frankel. But before this can happen, she gets knocked unconscious at a photo shoot of the "Million Dollar Bra" at the Guggenheim, spinning her into a dream sequence. An angelic Dr. Frankel shows her how different her life would have been had she never worn braces. She's the "pretty sister," while Hilda is homely, Ignacio is a compulsive gambler, Marc is a doting dad and Amanda is married to Daniel. But glam Betty is also mean - she's in cahoots with Wilhemina to terrify and overpower everyone at Mode.