Betty is thrilled to get her braces off, courtesy of her chatty new orthodontist, Dr. Frankel. But before this can happen, she gets knocked unconscious at a photo shoot of the "Million Dollar Bra" at the Guggenheim, spinning her into a dream sequence. An angelic Dr. Frankel shows her how different her life would have been had she never worn braces. She's the "pretty sister," while Hilda is homely, Ignacio is a compulsive gambler, Marc is a doting dad and Amanda is married to Daniel. But glam Betty is also mean - she's in cahoots with Wilhemina to terrify and overpower everyone at Mode.

As Hilda's wedding day approaches, Betty asks Daniel to be her plus-one, Ignacio invites Elena, and Amanda is set on fixing up Marc with her new client, soap opera star Spencer Cannon. Meanwhile, Justin weighs whether his own plus-one could be Austin, Betty considers unexpected offers in her personal and professional life, and Wilhelmina finds a way to fight the Meade empire by using Tyler as her pawn.