The Wielders find the Fool's Stone, and can now use their powerful Ignite modules against their alchemist foes. Cagliostro is the first to challenge them and their new power.

With what they've learned from the shrine, the Symphogear Wielders now know the full extent of the Illumanti's plan: they're going to use an ancient gate to summon the power of a god. Only one person can stop them: Hibiki, the "God-Killer".