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While Sam reels from the events of Waltringham and Joe struggles to help him through it, truths are revealed and dangerous plans put into action at the Railyard.
Laura leads a tense Dryden Commission on a visit to the Railyard, but will Agnes' anger threaten its success? Mia comes face to face with Ed forcing her to confront her past, while an evening out for Karen, Sam and Joe turns to tragedy.
Stanley and Anatole's true colours put Laura and her family directly in harm's way. With Mia speaking at the commission, Laura must do everything in her power to stop an awful attack. Leo and Mattie return to the Elster house where Leo comes to understand a painful truth, but Mattie has a secret that could knock their fledging relationship off course. Niska continues her mysterious journey that takes her to Wales, where she must defy all her instincts to survive.
One year after the dawn of consciousness, a decimated and oppressed Synth population fights to survive in a world that hates and fears them. In a divided Britain, Synths and Humans struggle to broker an uneasy peace, but when fractures within the Synth community itself start to appear, all hope of stability is threatened.Set against the maelstrom of political chaos, the ethical complexities of the dawn of a new species play out across a thrilling multi-stranded narrative. As the synth family, Mia, Niska and Max continue to battle for their right to survival, Joe, Laura and their children Mattie, Toby and Sophie struggle to come to terms with the cataclysmic events of the last series and their deeply polarised views on the implications of the singularity. Joining the cast this series is Mark Bonnar who plays Neil Sommer, an intriguingly charming scientist on a government commission with whom Laura develops a strong personal connection.