Broadway star Matthew Broderick knew little about his grandfather, James Joseph Broderick, and is shocked to learn about his involvement as a medic in World War I. Continuing his research, he discovers his grandmother, Mary Martindale, was a descendant from a Civil War hero, Robert Martindale, who fought at Gettysburg and joined the historic march through Georgia. While in Atlanta, Georgia, researching Robert's death, he unwittingly helps historians solve a 150 year-old mystery.

Academy Award winner Susan Sarandon has always been intrigued by her enigmatic grandmother, Anita Rigali, who abandoned her daughter, Leonora, when she was just two years old. Susan had grown up feeling a strange connection to her 'colorful' grandmother and had always wanted to solve the mystery of her life. With some clever sleuthing, Susan traces Anita's roots to Italy where she tracks down the actual village in Tuscany that was home to Anita's father. With the help of her son, Susan turns gumshoe and uncovers Anita's obituary, revealing the shocking missing link.