World-renowned Chef Jet Tila helps Tyler Florence challenge the five teams to make dishes that reflect their culinary origin stories -- and his favorite gets a reward! The pro versus rookie rivalry intensifies with catastrophic consequences for one team in particular. The rookies make huge strides, and in a shocking elimination, one of the top-ranked teams tumbles to the bottom two.

With a key team member out sick, one food truck is forced to operate as a two-person unit, placing them at a severe disadvantage. Culinary powerhouse Chef Joe Sasto joins Tyler Florence to challenge the final three teams to a taste and speed challenge. On day two, the competitors work beside some of Los Angeles' best food trucks for a nighttime rally in Venice Beach, where they'll meet Daniel Shemtob, head chef of two-time Race winner Lime Truck.