Jeremy tests the new Mitsubishi Evo X against the new Subaru Impreza WRX STi on the Top Gear track; Richard races the new Audi estate against two skiers in the Alps and Jeremy road-tests the new Mercedes CLK Black ("Imagine drinking a pint of hemlock, setting yourself on fire and then jumping out of a plane when it's directly overhead a combine harvester… That's what the Black feels like. Exciting beyond words. … I'm not sure that anyone's life is quite complete unless they have one"). Actors Peter Firth and Rupert Penry-Jones (famous for playing spies in the BBC drama series Spooks) are the "Stars in a Reasonably Priced Car."

In a race across Japan, Jeremy Clarkson tries to prove that the technical tour de force that is the Nissan GT-R is faster than Richard Hammond and James May traveling on the ultra-efficient Japanese public transport, including the 200 mph Shinkansen Bullet Train. Jeremy also tests the beautiful new Alpha Romeo 8C on the Top Gear track. UK newsreaders Fiona Bruce and Kate Silverton are the "Stars in a Reasonably Priced Car."