After their home collapses, two young squirrel brothers find their survival skills tested on a journey through a tree trunk towards freedom, meeting rodent friends and foes along the way. A snake and her mouse prey face off. Puppies at a pet store settle a turf war.

After his wife is taken from him, a turkey seeks revenge on The Mayor during the Thanksgiving parade. Two rat teens venture out of the sewer and discover the joys and horrors of life aboveground.