When the local Lark Rise gossip Mrs Mullins discovers her daughter Emily is pregnant, she leaps to the conclusion that Daniel is the father. Everyone, including Laura, rallies round to defend him. But when Daniel's behaviour starts to arouse suspicion, and people begin to doubt him, Dorcas is determined to uncover the truth. Can Laura and Daniel's relationship survive the crisis?

When the postmaster at Inglestone, with an old score to settle, tries to force Dorcas into selling up, she is faced with the heartbreaking prospect of losing her home and denying Sydney his dream of running the post office one day. But can Dorcas, with the help and determination of her staff and her friends, find a way to save the post office and their jobs? And with Laura torn between the love of two men, Daniel and Fisher, who will she choose?