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Jiro still tries his best to prove that he is stronger that Taro, while a new enemy appears before the Donbrothers.
One year has passed since Tsubasa made his promise to Natsumi and Tsubasa believes that soon he will reunite with her, but instead, he meets a young heartbroken man who is about to turn into a Hitotsuki.
Haruka meets a ghost who can't get to the afterlife because she is waiting for a call from the person she loves, while the Donbrothers face an enemy who has no corporeal form.
A man called Jin Momoi finds a baby inside a peach-shaped capsule whom he names Taro and raises. 21 years later, mysterious monsters called the Hitotsuki begin appearing when humans overcome with their inner desires are taken over by spirits infused with powers related to the Super Sentai from the past. Taro, and four other individuals selected by Jin; Shinichi Saruhara, a smart but eccentric young man, Haruka Kito, a highschooler manga artist, Tsubasa Inuzuka, a wanted fugitive, and Tsuyoshi Kijino, an ordinary salaryman begin fighting the Hitotsuki and saving their hosts as the Donbrothers.
Also standing on their way are the Noto, elusive and ruthless creatures from a higher plane who destroy the Hitotsuki while killing their hosts, unlike the Donbrothers who use their powers to rescue them. The Donbrothers also count with the secret help of Kaito Goshikida, the owner of the coffee shop Donbura who transforms into Zenkaizer Black