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Sonoshi, an inspector sent by the Noto Council, reprimands Sonoi, Sononi and Sonoza for getting distracted from their job and orders them to destroy the Donbrothers for good.
Tsubasa's search for Natsumi leads him to the Juto Forest, only to be captured with a Juto assuming his appearance. The Juto posing as Tsubasa is tailed by an unaware Tsuyoshi while Haruka meets Miho as she was acting on Natsumi's memories. Taro eventually confronts the two Juto prior to Sonoza intervening.
Having settled their dispute, Tsubasa and Tsuyoshi have a diner at a restaurant to celebrate, but the chef gets on an argument with Tsubasa and they decide to have a cooking competition.
A man called Jin Momoi finds a baby inside a peach-shaped capsule whom he names Taro and raises. 21 years later, mysterious monsters called the Hitotsuki begin appearing when humans overcome with their inner desires are taken over by spirits infused with powers related to the Super Sentai from the past. Taro, and four other individuals selected by Jin; Shinichi Saruhara, a smart but eccentric young man, Haruka Kito, a highschooler manga artist, Tsubasa Inuzuka, a wanted fugitive, and Tsuyoshi Kijino, an ordinary salaryman begin fighting the Hitotsuki and saving their hosts as the Donbrothers.
Also standing on their way are the Noto, elusive and ruthless creatures from a higher plane who destroy the Hitotsuki while killing their hosts, unlike the Donbrothers who use their powers to rescue them. The Donbrothers also count with the secret help of Kaito Goshikida, the owner of the coffee shop Donbura who transforms into Zenkaizer Black