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Haruka's aunt is kidnapped and used as a hostage by the Noto who force the Donbrothers to join them in their mission to hunt down the Juto.
Taro receives a funeral notice for Sonoi from the Noto Council. Taro, who decides to go alone, is invited to a different dimension, where he is surrounded by celestial maidens and receives hospitality such as baths, massages, and meals. Meanwhile, Don Murasame appears to help the Anoni, who is attacked by the Juto. Jiro tries to fight back, but the other Jiro and Murasame feel similar to each other.
The other Donbrothers finally discover that Tsubasa is Inu Brother, which brings him en route of collision with Tsuyoshi due to the connection between Miho and Natsumi.
A man called Jin Momoi finds a baby inside a peach-shaped capsule whom he names Taro and raises. 21 years later, mysterious monsters called the Hitotsuki begin appearing when humans overcome with their inner desires are taken over by spirits infused with powers related to the Super Sentai from the past. Taro, and four other individuals selected by Jin; Shinichi Saruhara, a smart but eccentric young man, Haruka Kito, a highschooler manga artist, Tsubasa Inuzuka, a wanted fugitive, and Tsuyoshi Kijino, an ordinary salaryman begin fighting the Hitotsuki and saving their hosts as the Donbrothers.
Also standing on their way are the Noto, elusive and ruthless creatures from a higher plane who destroy the Hitotsuki while killing their hosts, unlike the Donbrothers who use their powers to rescue them. The Donbrothers also count with the secret help of Kaito Goshikida, the owner of the coffee shop Donbura who transforms into Zenkaizer Black