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Haruka and the others look for a way to bring Taro back, while Jiro Momotani, another individual whose dream is to become a hero, appears before them.
Fed up with Taro's behavior, the other Donbrothers decide to fight the Hitostsuki without following his orders. Meanwhile, Taro and Sonoi have a duel after discovering each other's true identities.
While Haruka and Shinichi are crippled from their failed attempt to revive Taro and Tsuyoshi transforms into a Hitotsuki, Jiro obtains the power to transform into a Donbrother as well to help set things straight.
A man called Jin Momoi finds a baby inside a peach-shaped capsule whom he names Taro and raises. 21 years later, mysterious monsters called the Hitotsuki begin appearing when humans overcome with their inner desires are taken over by spirits infused with powers related to the Super Sentai from the past. Taro, and four other individuals selected by Jin; Shinichi Saruhara, a smart but eccentric young man, Haruka Kito, a highschooler manga artist, Tsubasa Inuzuka, a wanted fugitive, and Tsuyoshi Kijino, an ordinary salaryman begin fighting the Hitotsuki and saving their hosts as the Donbrothers.
Also standing on their way are the Noto, elusive and ruthless creatures from a higher plane who destroy the Hitotsuki while killing their hosts, unlike the Donbrothers who use their powers to rescue them. The Donbrothers also count with the secret help of Kaito Goshikida, the owner of the coffee shop Donbura who transforms into Zenkaizer Black