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The Donbrothers find themselves in great danger when they accidentally summon a Noto android called Don Killer who proves himself as their most powerful foe so far.
Having settled their dispute, Tsubasa and Tsuyoshi have a diner at a restaurant to celebrate, but the chef gets on an argument with Tsubasa and they decide to have a cooking competition.
Haruka is in trouble trying to get her driving license, while Don Murasame attacks Tsubasa who is saved in the nick of time by Sononi, who stands up to protect Tsubasa out of her feelings for him.
A man called Jin Momoi finds a baby inside a peach-shaped capsule whom he names Taro and raises. 21 years later, mysterious monsters called the Hitotsuki begin appearing when humans overcome with their inner desires are taken over by spirits infused with powers related to the Super Sentai from the past. Taro, and four other individuals selected by Jin; Shinichi Saruhara, a smart but eccentric young man, Haruka Kito, a highschooler manga artist, Tsubasa Inuzuka, a wanted fugitive, and Tsuyoshi Kijino, an ordinary salaryman begin fighting the Hitotsuki and saving their hosts as the Donbrothers.
Also standing on their way are the Noto, elusive and ruthless creatures from a higher plane who destroy the Hitotsuki while killing their hosts, unlike the Donbrothers who use their powers to rescue them. The Donbrothers also count with the secret help of Kaito Goshikida, the owner of the coffee shop Donbura who transforms into Zenkaizer Black