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John Craven and Sammi Kinghorn visit the South Downs to see how the past is inspiring residents to preserve the future of this diverse landscape.
As part of Countryfile's Wild Britain initiative, Ellie Harrison and Hamza Yassin visit Wytham Woods in Oxfordshire.
Countryfile returns to Dumfries House as autumn is in full swing and the first year of teaching gets underway at the new rural skills centre recently opened by the King.