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Maki and Kazari experiment with Gamel in determining the use of a Greeed for a Core Medal container as Eiji approaches Kougami in finding out more regarding the artificial beings.
Kazari uses a transport truck of Cell Medals, a Core Medal of Ankh and a captured Hina to lure all sides into a free-for-all showdown.
Eiji and Ankh try to figure out Akira Date, the man who has become Kamen Rider Birth, as Uva creates a peculiar Yummy from a kendo girl who wishes to be stronger.
Eiji Hino is a traveling man who has no place to call home and a tragic past. When coin-based homunculi known as the Greeed awaken after their 800-year slumber to attack humans and feed off of their desires, the disembodied arm of the Greeed named Ankh gives Eiji a belt and three Medals to fight the other Greeed as Kamen Rider OOO. The mysterious Kougami Foundation approaches Eiji and begins assisting him in his fight against the Greeed, though their true motives are not clear. As Eiji fights the Greeed and their Yummy monsters, learning more of the Greeed and Ankh, he starts to find a purpose beyond his journey.