When devastating truths behind the Mars attack are revealed, Picard realizes just how far many will go to preserve secrets stretching back generations, all while the La Sirena crew grapples with secrets and revelations of their own. Narissa directs her guards to capture Elnor, setting off an unexpected chain of events on the Borg cube.

A final confrontation on the synthetics' homeworld, Coppelius, pits Picard and his team against the Romulans, as well as the synths who seek to safeguard their existence at all costs.