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Joe continues her training with Cruz; her methods are questionable. Cruz starts to build a relationship with Aaliyah. Kate gets into a fight during her soccer game. Neal has to deliver difficult news to a patient.
Things go awry for Joe and her team during a mission out in the field; Joe is left devastated. Upon her return home, Joe's family life presents it's own challenges. Cruz tries to escape a dangerous situation. Joe must choose a new recruit.
Cruz is invited to spend the weekend with Aaliyah and her inner-circle. Joe keeps Kaitlyn informed of Cruz's progress. Kyle comes to Joe desperate for help with an asset.
Special Ops: Lioness, inspired by an actual U.S. Military program, follows the life of Joe while she attempts to balance her personal and professional life as the tip of the CIA's spear in the war on terror. The Lioness Program, overseen by Kaitlyn Meade and Donald Westfield, enlists an aggressive Marine Raider named Cruz to operate undercover alongside Joe among the power brokers of State terrorism in the CIA's efforts to thwart the next 9/11.