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Echo and Victor's imprints collide when Echo is sent on an assignment as a fun-loving, seductive college student and Victor is imprinted as the psychotic nephew of a Dollhouse shareholder.
When Echo is imprinted as a mother with a newborn baby, she takes too strongly to motherhood as a result of Topher's modifications. Meanwhile, Adelle pays November a visit, and Perrin ramps up his investigation into the Rossum Corporation.
Sierra is forced to face her past as her connection to the Rossum Corporation leads to obsession and murder.
The Dollhouse is a very secret, and very illegal, place where wishes come true. Clients with the right connections and enough money can hire "Actives", people who have been programmed to perfectly fulfill the needs, and desires of their clients. The Actives are people who have chosen, for their own reasons, to surrender their bodies and minds for a five-year stay in the Dollhouse. Now they can be imprinted with any personality, skill, or even muscle memory. They can be the perfect companion, lover, spy, assassin; and when the job is done they forget everything. But something is wrong with the Active called "Echo". No longer just a blank slate waiting for her next assignment, Echo is remembering flashes of the lives that she has lived and the games she has played, and she is starting to wonder just who she really is.