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In the wake of Jasper's actions against the Maida Vale, police take suspects into custody. Meanwhile, Helen makes a significant discovery about her own past.
Guy and Jasper have a head-to-head meeting at the London Motherhouse, and Helen's true motives are questioned.
Guy breaks away from his identity and has a confrontation with Jasper, who sends him on a mission to a vampire coven's hotel to find the "Seven Five Two".
After a dangerous mission, Guy's trust in Helen and the Talamasca is shaken. Helen recounts the story of the "Seven Five Two," revealing a key part of the organization's history.
Guy joins the Talamasca, leaving his old life as a lawyer behind. Helen trains him on the ins and outs of undercover work. Guy is sent to London on his first mission – to follow a mysterious hit man.
Guy is recruited to join a top-secret spy organization for the supernatural world; he must choose between his old life in the ordinary world and a new life as an undercover spy; Helen deals with trouble at the New York Mother House.