Lucie and Charles journey to Paris in search of her long-lost father. The two grow closer, until Charles' past rears its head, back in London, Lucie is drawn to the troubled Sydney Carton. Thérèse Defarge plots revenge against the Marquis.

As the Revolution descends into bloodshed, Charles is arrested in Paris; desperate to save him, Lucie turns to Sydney Carton, but at Charles' trial, a revelation changes everything; as Thérèse seeks vengeance, Sydney weighs up an epic sacrifice.