Leopard mother Olimba is bringing up her two cubs in an ideal leopard territory called Nsefu, in Zambia. Soon, however, her world is turned upside down by the arrival of wild dog Storm and her chaotic pack. Storm's pack seems unchallengeable – even dominating hyena Tandala and the rest of her clan. But just when the dogs are beginning to settle into the kingdom and welcome a litter of puppies, an even more powerful force appears on the horizon – lions. These new arrivals are a threat to everyone here – something that is brought into sharp focus when Olimba's young son goes missing...

In a land of extreme seasons and intense rivalry, motherhood is the greatest trial females in Nsefu face. Lioness Rita is raising the next generation, but caring for her cubs alone leaves them vulnerable while she hunts. The hyenas' strength is growing, but a flooded den spells disaster, and the clan become separated. Their leader Tenta must protect her twins as they navigate the watery labyrinth to find the rest of their scattered family. Seeking a mate, leopard Olimba begins a new chapter, but in her way is her daughter – she has now become her main rival, and only one can remain...