At Britain's oldest public museum, the Ashmolean Museum of Art and Archaeology in Oxford, being put to the test by Griff are Lars Tharp, Professor Kate Williams, Dr Janina Ramirez and Dr Michael Scott. The two teams pit their knowledgeable wits against each other in a series of rounds informed by the amazing contents of the museum.

At Queen's House at the National Maritime Museum in Greenwich, the experts being put to the test by Griff Rhys Jones are historian Professor Kate Williams, historian specialising in European and Chinese ceramics Lars Tharp, former first sea lord Admiral Lord West, and physicist and oceanographer Helen Czerski.